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A young Carl Fredrickson meets a young adventure spirited girl named Ellie. They both dream of going to a Lost Land in South America. 70 years later, Ellie has died. Carl remembers the promise he made to her. Then, when he inadvertently hits a construction worker, he is forced to go to a retirement home. But before they can take him, he and his house fly away. However he has a stowaway aboard. An 8 year old boy named Russell, who's trying to get an Assisting the Elderly badge. Together, they embark on an adventure, where they encounter talking dogs, an evil villain and a rare bird named Kevin.

Carl Fredrickson, a little boy and a dreamer who idolizes the adventurer Charles Munts. When he meets Ellie, who also worships Munts, they become close friends. However Charles Munts falls into disgrace, accused of forging the skeleton of the monster of Paradise Falls. He travels in his blimp to South America to bring the monster back alive but is never seen again. Eventually he Carl grows up and marries Ellie. They promise each other that they would travel together to Paradise Falls and build a house there. Many years later, Ellie dies and Carl, who's lonely, refuses to move from their house despite the offers of the owner of a construction company. When Carl accidentally hits a worker that damaged his mailbox, he is sentenced to move to a retirement home. However, he uses many balloons to float his house in order to travel to Paradise Falls. Adventure ensues.